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Pickguard off? Covers off
On the original question, I'm going to hedge my bets... if the guitar is vintage-style, a bit mellowed-looking, nickel hardware etc I'd keep the covers on. If it's new and shiny with chrome hardware I'd take the covers off.
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Sound is far more important though. They will sound slightly different with or without the covers. Generally slightly less high end with the covers.
Black LP custom - either
LP Standard - on
SG - deffo on
Superstrat - deffo off
PRS - off on 24 fret models on on 22